Editorial Board:
Chair: Professor John Coggins, OBE, FRSE, FSB
Emeritus Professor of Molecular Enzymology, School of Biology,
University of Glasgow
Professor Peter Grant, OBE, FRSE, FIET, FIEE, FREng
Emeritus Regius Professor of Engineering, School of Engineering, University of Edinburgh
Professor Angus Lamond, FRS, FRSE
Professor of Biochemistry, University of Dundee
Professor David Milne, OBE, FRSE, FIET, SMIEEE, FREng
Non-Executive Director, Wolfson Microelectronics plc
Professor Paul Bishop, FRSE, Fellow of the Geological Society of America,
Professor of Geography, School of Geography & Earth Sciences, University of Glasgow
Editorial Team:
Writer: Peter Barr
Designer: Emma Quinn
Production Editor: Jenny Liddell, The Royal Society of Edinburgh
Printer: Mackay & Inglis Ltd
www.royalsoced.org.uk
www.sciencescotland.org
ISSN 1743 – 6222 (Science Scotland Print)
ISSN 1743 – 6230 (online)
If you would like more information, please contact: sciencescotland@royalsoced.org.uk
The Royal Society of Edinburgh, 22–26 George Street, Edinburgh EH2 2PQ
ANY VIEWS EXPRESSED IN THIS PUBLICATION DO NOT NECESSARILY REPRESENT THOSE
OF THE ROYAL SOCIETY OF EDINBURGH, NOR ALL OF ITS FELLOWS.
The Royal Society of Edinburgh, Scotland's National Academy, is Scottish Charity No SC000470